By virtue of the alpaca database held at PACOMARCA, the world's widest and most complete, and the collaboration with the Complutense University of Madrid, many scientific papers have been written and published in the most important international indexed journals.

PACOMARCA also frequently presents non-scientific information on its genetic-improvement program in different international and national publications as well as at veterinary conferences, meetings of groups brought together by a common interest in alpaca, specialized events, university-level lectures and so on.

  • INDEXED JOURNALS

    CRUZ, A., MURILLO, Y., BURGOS, A., YUCRA, A., MORANTE, R., QUISPE, M., QUISPE, C., QUISPE, E., GUTIERREZ, J.P. 2024. Genetic parameters for different types of medullated fibre in Alpacas. Journal of Animal Breeding and Genetics. Online ahead of print.
    MORE, M., VELI, E., CRUZ, A., GUTIÉRREZ, J.P., GUTIÉRREZ, G., PONCE DE LEÓN BRAVO, F.A. 2023. Genome-Wide Association Study of Fiber Diameter in Alpacas. Animals 13, 3316.
    QUISPE, E., TORRES, R., PINARES, R., CRUZ, A., QUISPE, M.D. 2023. Comparison of projection microscope with OFDA100 in alpaca fibers medullation. Archivos de Zootecnia 72, 226-234
    GUTIÉRREZ, J.P., CRUZ, A., MORANTE, R., BURGOS, A., FORMOSO-RAFFERTY, N., CERVANTES, I. 2023. Genetic parameters for fleece uniformity in Alpacas. Journal of Animal Science 101.
    CRUZ, A., SEDANO J., BURGOS A., GUTIÉRREZ J. P., WURZINGER M. AND GUTIÉRREZ-REYNOSO G. 2023. Genomic selection improves genetic gain for fiber traits in a breeding program for alpacas. Livestock Science 270. doi:10.1016/j.livsci.2023.105195.
    QUISPE, E., QUISPE, M.D., QUISPE, C., POMA, A., PAUCAR-CHANCA, R., CRUZ, A., McGREGOR, B. 2022. Relationship between the incidence and degree of medullation with the diameter of alpaca fibers evaluated using a novel device based on artificial intelligence. The Journal of the Textile Institute 1-16.
    QUISPE, M.D., SERRANO-ARRIEZU, L., TRIGO, J.D., QUISPE, C., POMA, A., QUISPE, E., 2022. Application of artificial intelligence and digital images analysis to automatically determine the percentage of fiber medullation in alpaca fleece samples. Small Rumin Res 213, 106724.
    MANCISIDOR, B., CRUZ, A., GUTIÉRREZ, G., BURGOS, A., MORON, J., WURZINGER, M., GUTIÉRREZ, J.P. 2021. ssGBLUP method improve the accuracy of breeding value prediction in Huacaya alpaca. Animals. 11 (11), 3052. doi:10.3390/ani11113052.
    CRUZ, A., GUTIÉRREZ, G.A., BURGOS, A., MORANTE, R., WURZINGER, M., GUTIÉRREZ, J.P. 2021. Software de gestión para pedigrí y producción de camélidos de nuevo mundo: Pacokipu y Llamakipu. Rev. Inv. Vet. Perú 32(4), e19355.
    CRUZ, A., YUCRA, A., GUTIÉRREZ, G.A., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., WURZINGER, M. 2021. Colorimetry analysis of coat color and its relationship with fiber traits in alpacas. Animal 15 (5) 100219.
    PINARES, R., CRUZ, A., DAVERIO, M.S., GUTIÉRREZ, J.P., PONCE DE LEÓN, F.A., WURZINGER, M., Di ROCCO, F., GUTIÉRREZ, G.A. 2021. Polimorfismos de nucleótido simple (PNSs) del gen MC1R en alpacas negras y marrones. Revista Peruana de Biología, 28(1), e19742. doi:10.15381/rpb.v28i1.19742 (Only Spanish)
    MARCOS CALDERON 1,2, MANUEL J. MORE 1, GUSTAVO A. GUTIÉRREZ 1 AND F. ABEL PONCE DE LEÓN 3,* 2021. Development of a 76k Alpaca (Vicugna pacos) Single Nucleotide Polymorphisms (SNPs) Microarray. MDPI
    CRUZ, A., GUTIÉRREZ, J.P., TORRES, R., FORMOSO-RAFFERTY, N., MORANTE, R., BURGOS, A., CERVANTES, I. 2020. Calving date and its variability as a potential trait in the breeding objective to account for reproductive seasonality in alpacas. Reprod Domest Anim. 2020;00:1–8.
    CRUZ, A., MENÉNDEZ-BUXADERA, A., GUTIÉRREZ, G., MORANTE, R., BURGOS, A., GUTIÉRREZ, J.P. 2020. Genetic (co) variance across age of fiber diameter and standard deviation in Huacaya alpacas, estimated by repeatability, multi-trait and random regression models. Livestock Science 231,103863.
    PINARES, R., GUTIÉRREZ, G.A., CRUZ, A. BURGOS, A., GUTIÉRREZ, J.P. 2019. Phenotypic variability of percentage of medullated fibers in Huacaya alpaca fleece. Rev. Inv. Vet. Perú 30, 699-708.
    CRUZ, A., MORANTE, R., GUTIÉRREZ, TORRES, R. ,J.P., BURGOS, A., CERVANTES, I. 2019. Genetic parameters for medullated fiber and its relationship with other productive traits in alpacas. Animal 13, 1358-1364.
    PINARES, R., GUTIÉRREZ, G., CRUZ, A., MORANTE, R., CERVANTES, I., BURGOS, A., GUTIÉRREZ, J.P. 2018. Heritability of individual fiber medullation in Peruvian alpacas. Small Ruminant Research 165, 93-100.
    CRUZ, A., CERVANTES, I., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., 2017. Genetic parameters estimation for preweaning traits and their relationship with reproductive, productive and morphological traits in alpaca. Animal 11, 746-754.
    CRUZ, A., MORANTE, R., CERVANTES, I., BURGOS, A., GUTIÉRREZ, J.P., 2017. Effect of the gestation and lactation on fiber diameter and its variability in Peruvian alpacas. Livestock Science 198, 31-36.
    CRUZ, A., CERVANTES, I., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., 2015. Estimation of genetic parameters for reproductive traits in alpacas. Animal Reproduction Science 163, 48-55.
    GUTIÉRREZ, J.P., CERVANTES, I., PÉREZ-CABAL, M.A., BURGOS, A., MORANTE, R., 2014. Weighting fibre and morphological traits in a genetic index for an alpaca breeding program. Animal 8, 360-369.
    PAREDES, M.M., MEMBRILLO, A., GUTIÉRREZ, J.P., CERVANTES, I., AZOR, P.J., MORANTE, R., ALONSO-MORAGA, A., MOLINA, A., MUÑOZ-SERRANO, A., 2014. Association of microsatellite markers with fiber diameter trait in Peruvian alpacas (Vicugna pacos). Livestock Science 161, 6-16.
    GUTIÉRREZ, J.P., VARONA, L., PUN, A., MORANTE, R., BURGOS, A., CERVANTES, I., PÉREZ-CABAL, M.A., 2011. Genetic parameters for growth of fiber diameter in alpacas. Journal of Animal Science 89, 2310-2315.
    CERVANTES, I., PEREZ-CABAL, M.A., MORANTE, R., BURGOS, A., SALGADO, C., NIETO, B., GOYACHE, F., GUTIÉRREZ, J.P., 2010. Genetic parameters and relationship between fibre and type traits in two breeds of Peruvian alpacas. Small Ruminant Research 88, 6-11.
    PÉREZ-CABAL, M.A., CERVANTES, I., MORANTE, R., BURGOS, A., GOYACHE, F., GUTIÉRREZ, J.P., 2010. Analysis of the existence of major genes affecting alpaca fiber traits. Journal of Animal Science 88, 3783-3788.
    GUTIÉRREZ, J.P., GOYACHE, F., BURGOS, A., CERVANTES, I., 2009. Genetic analysis of six production traits in Peruvian alpacas. Livestock Science 123, 193-197.
    MORANTE, R., GOYACHE, F., BURGOS, A., CERVANTES, I., PÉREZ-CABAL, M.A., GUTIÉRREZ, J.P., 2009. Genetic Improvement for alpaca fibre production in the Peruvian altiplano: The Pacomarca experience. Animal Genetic Resources Information 45, 37-43.
    PINARES, R., CRUZ, A., MORANTE, R., CERVANTES, I., BURGOS, A., GUTIÉRREZ, G., GUTIÉRREZ, J.P. 2019. Heredabilidad estimada de fibras meduladas en alpaca huacaya. Universitätsverlag Göttingen. Advances in Fibre Production Science in South American Camelids and other Fibre Animals. Chapter: Breeding and Genetics. pp 77-82.
    MUNYARD, k., GREEFF, J. 2013. Quantitative genetic analysis of micron blowout in alpacas. Australian Government, Rural Industries Research and Development Corporation.RIRDC PRJ-006076, 12/136.
  • OTHER PUBLICATIONS

    BURGOS, A., GUTIÉRREZ, J.P., CRUZ, A., GUTIÉRREZ, G. 2023. PACOMARCA, Alpacas: genética en blanco y negro. 1ra edición, Lima, Perú. ISBN:978-612-49369-0-6.
    CRUZ, A., MURILLO, Y., BURGOS, A., YUCRA, A., QUISPE, M.D., QUISPE, E., GUTIÉRREZ, J.P. 2023. Genetic parameters of medullation types in alpaca fiber. 74nd Annual Meeting of the European Federation of Animal Science: EAAP + WAAP + Interbull Congress 2023. August 26th to September 1st. Lyon, France. Abstract Nro. 41503.
    GUTIÉRREZ R, G., CRUZ, A., WURZINGER, M., GUTIÉRREZ, J.P. 2023. La genómica al rescate de la fibra de alpaca. Hominem Et Agrum, 1(1), 25-26. Revista de la Universidad Nacional Agraria La Molina.
    NÖLLENBURG, N., CRUZ A., BURGOS, A., GUTIÉRREZ, G., WURZINGER, M., WINCKLER, C. 2022. Stress in alpacas: physiological and behavioral responses during the shearing procedure. 8thEuropean Symposium on South American Camelids & 4th European Meeting on Fibre Animals. September 26 to 28, Bolzano, Italy.
    CRUZ, A., GUTIÉRREZ, G., YUCRA, A., BURGOS, A., WURZINGER, M., GUTIÉRREZ, J.P. 2022. Improving a black alpaca breeding program using a colorimeter values for male selection. 8th European Symposium on South American Camelids & 4th European Meeting on Fibre Animals. September 26 to 28, Bolzano, Italy.
    YUCRA, A., BURGOS, A., GUTIÉRREZ, G., WURZINGER, M., GUTIERREZ, J.P., CERVANTES, I., CRUZ, A. 2022. Relationship between coat colour and temperature measurements along the alpaca fibre. 8thEuropean Symposium on South American Camelids & 4th European Meeting on Fibre Animals. September 26 to 28, Bolzano, Italy.
    PINARES, R., CRUZ, A., DAVERIO, M.S., DI ROCCO, F., GUTIERREZ, G. 2022. Polymorphisms in MC1R and ASIP genes associated with color phenotypes in alpaca Huacaya. 8thEuropean Symposium on South American Camelids & 4th European Meeting on Fibre Animals. September 26 to 28, Bolzano, Italy.
    GUTIÉRREZ, G., CRUZ, A., GUTIÉRREZ, J.P., WURZINGER, M. 2021. Genetic improvement of alpacas and llamas in Peru. 72nd Annual Meeting of the European Federation of Animal Science. August 30 to September 3. Davos, Switzerland. pp.292.
    WURZINGER, M., SILVA, C., CANDIO, J., CRUZ, A., QUINA, E., GUTIÉRREZ, G.A. 2021. Attitudes of Peruvian llama farmers towards genetic improvement. 72nd Annual Meeting of the European Federation of Animal Science. August 30 to September 3. Davos, Switzerland. pp.293.
    MANCISIDOR, B., CRUZ, A., GUTIÉRREZ, G.A., BURGOS, A., MORON, J., WURZINGER, M., GUTIÉRREZ, J.P. 2021. Efficiency of genomic selection for fiber traits in alpacas. 72nd Annual Meeting of the European Federation of Animal Science. August 30 to September 3. Davos, Switzerland. pp.446.
    MANCISIDOR, B., CRUZ, A., GUTIÉRREZ, G.A., BURGOS, A., MORON, J., WURZINGER, M., GUTIÉRREZ, J.P. 2021. Comparison of breeding values accuracy using BLUP and ss-GBLUP methodology in Peruvian alpacas. International symposium on sustainable animal production and health – Current status and way forward. Jun 28 to july 2. Virtual meeting. pp.428-431.
    CRUZ, A., QUISPE-POTOSINO, S., CERVANTES, I., BURGOS, A., GUTIÉRREZ, J.P. 2020. Assessment of two shearing practices on animal welfare in alpaca using infrared thermography. Session 65, Poster 10, Abstract 33770. 71th Annual Meeting of European Federation for Animal Science. Virtual meeting. December 1st to 4th. pp. 588.
    CRUZ, A., CERVANTES, I., TORRES, R., FORMOSO-RAFFERTY, N., MORANTE, R., BURGOS, A., GUTIÉRREZ, J.P. 2019. Calving date as a potential breeding objective to manage the reproductive seasonality in alpacas. 70th Annual Meeting of the European Federation of Animal Science. 26-30 august. Ghent, Belgium.
    CRUZ, A., BURGOS, A., TORRES, R., GUTIÉRREZ, JP. 2018. Analysis of the body weight and prediction of the growth curve using non-linears models in Peruvian alpacas. VIII Congreso Mundial de Camélidos. Octubre 21 – 23, Oruro Bolivia, pp 56-57.
    GUTIÉRREZ, J.P. 2018. La médula como criterio de selección. VIII Congreso Mundial de Camélidos. Octubre 21 – 23, Oruro Bolivia, pp 28.
    GUTIÉRREZ, G., GUTIÉRREZ, J.P., HUANCA, T., WURZINGER, M., 2018. Challenges and opportunities of genetic improvement in alpacas and llamas in Peru. 11th World Congress of Genetics Applied to Livestock Production, 12-16 Febrero. Auckland, New Zealand.
    GUTIÉRREZ, J.P. 2017. Desarrollo de un programa de mejora de alpacas en Perú. 46th Congreso Argentino de Genética y IV Foro Regional del NOA. 1 al 4 de Octubre, 2017. SFV Catamarca. Argentina.
    CRUZ, A., MORANTE, R., CERVANTES, I., BURGOS, A., GUTIÉRREZ, J.P., 2017. Genetic parameters of medullation percentage in alpacas. 68th European Association for Animal Production (EAAP), 28 agosto-4 septiembre. Tallin. Estonia. pp. 369.
    FRANK, E.N., FRONDIZI SEGHETTI, D.G., HICK, M.V.H., CASTILLO, M.F., BURGOS, A., CRUZ, A. 2017. Dehairing of alpaca fibres top with AM2 dehairing technology. 7th European Symposium on South American Camelids and 3rd European Meeting on Fibre Animals. 12-14 de Junio, Assisi, Italy. pp. 71
    PINARES, R., CRUZ, A., MORANTE, R., CERVANTES, I., BURGOS, A., GUTIÉRREZ, G., GUTIÉRREZ, J.P. 2017. Heritability estimate of medullated fibre in Alpaca Huacaya. 7th European Symposium on South American Camelids and 3rd European Meeting on Fibre Animals. 12-14 de Junio, Assisi, Italy. pp. 27.
    CRUZ, A., 2017. Parámetros genéticos de caracteres funcionales y secundarios en alpacas. Tesis doctoral, Universidad Complutense de Madrid, España.
    CRUZ, A., CERVANTES, I., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., 2016. Heritabilities for preweaning and its genetic correlations with other traits in Peruvian alpacas. 67th European Association for Animal Production (EAAP), 29 Agosto - 2 Septiembre. Belfast, United Kingdom. pp. 520.
    GUTIÉRREZ, J.P., 2015. Construcción de índices de selección para programas de mejora genética en alpacas, VII World congress on South American Camelids, 28-30 de octubre, Puno, Perú. pp. 248-253.
    CRUZ, A., BURGOS, A., MORANTE, R., CERVANTES, I., GUTIÉRREZ, J.P., 2015. Importancia de la estima de parámetros genéticos de caracteres productivos y funcionales en alpacas. VII World congress on South American Camelids, 28-30 de octubre. Puno. Perú. pp. 276-280.
    BURGOS, A., APAZA, N., TORRES, R., CRUZ, A., 2015. La “Inca Esquila”: Esquila tecnificada para la industria textil. VII World congress on South American Camelids. 28-30 de octubre, Puno, Perú. pp. 718-719.
    CRUZ, A., CERVANTES, I., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., 2015. Estimation of genetic parameters for reproductive traits in alpacas. 66th European Association for Animal Production (EAAP). 31 Agosto-4 Septiembre, Varsovia, Polonia.
    CRUZ, A., 2015. Resultados de la selección, en busca de la alpaca textil en Pacomarca. Foro Internacional sobre avances en la producción. Mejoramiento y transformación de fibras de llamas, alpacas y vicuñas. 16 al 18 de Julio, Abancay, Perú. pp. 67-71.
    GUTIÉRREZ, J.P., 2013. Fibre genetics on Alpaca. 6th Symposium on South American Camelids and 2nd European meeting on Fiber Animals, (Ed. by D. Allain), In: 64th EAAP Annual meeting. 25-30 August, Nantes, France.
    GUTIÉRREZ, J.P., CERVANTES, I., PÉREZ-CABAL, M.A., PUN, A., BURGOS, A., MORANTE, R., 2012. Economically weighting fibre and morphological traits in an alpaca breeding program. 63th European Association for Animal Production (EAAP). 27-31 Agosto, Bratislava, Slovakia.
    PUN, A., MORANTE, R., BURGOS, A., CERVANTES, I., PÉREZ-CABAL, M.A., GUTIÉRREZ, J.P., 2011. Genetic parameters for fiber diameter at different shearings. 62th European Association for Animal Production (EAAP). 29 agosto-2 septiembre, Stavanger, Noruega.
    GUTIÉRREZ, J.P., 2011. Animal Breeding in Alpaca Suri. 2011 Suri Network Summer Symposium. Chicago, 4 al 6 de agosto.
    MORANTE, R., BURGOS, A., GUTIÉRREZ, J.P., 2010. Producing alpaca fibre for the textile industry. 5th European Symposium on South American Camelids and First European Meeting on Fibre Animals. 6 al 8 de octubre, Sevilla, España.
    VARONA, L., CERVANTES, I., PÉREZ-CABAL, M.A., MORANTE, R., BURGOS, A., GUTIÉRREZ, J.P., 2010. Genetic parameters for growth of fibre diameter in alpacas. 5th European Symposium on South American Camelids and First European Meeting on Fibre Animals. 6 al 8 de octubre, Sevilla, España.
    MORANTE, R., BURGOS, A., GUTIERREZ, J.P., 2010. Mejoramiento Genético de fibra de alpaca para la industria textil. III Simposium Internacional de Investigaciones sobre Camélidos Sudamericanos. 8 al 10 de septiembre, Arequipa, Perú.
    GUTIÉRREZ, J.P., CERVANTES, I., PÉREZ-CABAL, M.A., MORANTE, R., BURGOS, A., SALGADO, C., NIETO, B., VARONA, L., GOYACHE, F., 2010. Resultados de experiencias de investigación sobre fibra de alpaca. III Simposium Internacional de Investigaciones sobre Camélidos Sudamericanos. 8 al 10 de septiembre, Arequipa, Perú
    PÉREZ-CABAL, M.A., CERVANTES, I., MORANTE, R., BURGOS, A., GOYACHE, F., GUTIÉRREZ, J.P., 2010. Genes mayores afectando al diámetro de fibra en alpacas. XV Reunión Nacional de Mejora Genética Animal. 16-18 de Junio, Vigo, Pontevedra
    CERVANTES, I., GOYACHE, F., PÉREZ-CABAL, M.A., NIETO, B., SALGADO, C., BURGOS, A., GUTIÉRREZ, J.P., 2009. Parámetros genéticos y tendencias genéticas asociados a caracteres productivos y de apariencia del vellón en alpacas. Zaragoza
    CRUZ, A., TORRES, R., BURGOS, A., GUTIÉRREZ, J.P., 2016. Influence of effect of pregnancy and lactation on alpaca fiber diameter in estimated breeding values. Proceedings of the VI Peruvian Congress Animal Reproduction of the Asociación Peruana de Reproducción Animal (ASPRA). August 17 to 20th, 2016, Arequipa, Peru. Spermova, 6(1): 62, pp. 108.
    GUTIERREZ J.P., 2010. Resultados de experiencias de investigacion sobre fibra de alpaca. Arequipa

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